Tribe
Anisodactylini
Dicheirus piceus. Photo: (c) Anna Caro, some rights reserved (CC BY-NC-ND)
Harpalinae is a huge subfamily of ground beetles that contains roughly 6,400 species worldwide. A rarely used common name for the subfamily is the harp beetles. The Harpalinae contain the most apomorphic ground beetles, displaying a wide range of forms and behaviors. Some are, rare among ground beetles, omnivores or even herbivores.